Koeppen, 1899
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Koeppen, N., 1899: Hyalosaccus ceratii nov. gen. et sp., parazit' dinoflagellat' Kievskogo Obshchestvo Estestvoispytateley, Zapiski, 16(1), p.89-135, pl. 6-8. (Cited as Koeppen, 1903 in J. Cachon, 1964.)